Page 4 - 《广西植物》2021年第12期
P. 4
ইᤩІᧇ̵ዛᧇዛইইইইইইইইইইই ᦵᓣዞॹ ԣԔԣϓই Ⴌᅳᤀᤫ ϓ ટᅳᤫ ϓԣ
ইইইইইইইইইইইইইইইইইইইইই ྉ᤹ᅳႥᡙ၊ᤀᡥɯ
ࣷ᧣ટ௴ጛટ௴
ᡥᡙᓣቂ ᐹႥʐ ጛצᅳᤀࣩᡙᔠᅳႥ
ۗᓣʐᔠዞࣩᤀᐹʗᔠ ᡓᐹႥʐᐹႥᔠᐹ ྉᨃʢᆍᤦᑕႿዹၤᑕዹᔀᑕᔀɯᖔ ᑕ Ⴟᔀऔ ༁ᡱᔀዹᔽᔀ༁ ᆍ ۗᓣʐᔠዞࣩᤀᐹʗᔠ ʢᆍቝ ᡷၤᔀ ۼᆍᆍႿ੫ શᑕᡷᔽᆍႿᑕ શᑕᡷࣼʢᔀ ᔀ༁ᔀʢᔀᖔ ୩ᔽዹၤࣼᑕႿ
জ ۪ʢᆍᔽႿዹᔀᖔ ऊၤᔽႿᑕ ҭҭҭҭҭҭҭҭҭ ᓂ᧧શ ̀ᆍႿ੫ीᔽᑕႿ੫ᖔ ऊ̀ጶશ᥈ ɟࣼᔀၤᆍႿ੫ᖔ ᓂ᧧В ᆍႿ੫ᖔ ɟ᧧શ ᥘᔽႿᖔ ɟВ ۼᔀႿᤦᔽႿྉϟँ࿗ँɯ
᥈ᔀႿᆍቝᔽዹ ᔽʛᔀႿᡷᔽᔽዹᑕᡷᔽᆍႿ ᑕႿʛ ዹၤᑕʢᑕዹᡷᔀʢᔽझᑕᡷᔽᆍႿ ᑕႿᑕᢃ༁ᔽ༁ ᆍ ᡷၤᔀ ᡱၤᆍ༁ᡱၤᑕᡷᔀ ᡷʢᑕႿ༁ᡱᆍʢᡷᔀʢ ᡱʢᆍᡷᔀᔽႿ ϟ ᑕቝᔽᢃ ੫ᔀႿᔀ ᔽႿ ᡱᔽႿᔀᑕᡱᡱᔀ ҭҭҭ
ҭҭҭҭҭҭҭҭҭҭҭҭҭҭҭҭҭ ̀ዶશ ᓂᔽၤᆍႿ੫ᖔ ᓂ᧧В ऊၤᑕᆍᖔ Ի̀ዶᨃ ᥘᔽႿ੫ᢃࣼᖔ ̀В ᓂᔽᢼࣼᑕႿᖔ ̀В ɟࣼ༁ၤࣼᑕႿ੫ྉϟँጢጢɯ
۪ၤᢃᆍ੫ᔀႿᔀᡷᔽዹ ᔀᆍࣼᡷᔽᆍႿ ᑕႿʛ ᔀᡱʢᔀ༁༁ᔽᆍႿ ᑕႿᑕᢃ༁ᔽ༁ ᆍ ዛۗᣏ ੫ᔀႿᔀ ᔽႿ ۗ၊ᡥᤀᤀᅳᡙᐹዞ၊ᡥ ᓣʐࣩᤀᔠ ҭҭҭҭҭҭҭҭҭҭҭҭҭҭҭ
ҭҭҭҭҭҭҭҭҭҭҭҭҭҭҭҭҭҭҭҭҭҭҭ Ի̀ዶશ᥈ શᔽᖔ ऊ̀ጶશ ᔀᆍႿ੫ᖔ ऊВ᧧ ାᆍᔽᑕႿ੫ᖔ ɟዶશ᥈ ɟᑕႿ੫ྉϟँᤃ࿗ɯ
୩ᔀᑕ༁ᆍႿᑕ ዹၤᑕႿ੫ᔀ༁ ᆍ ዹʢᢃ༁ᡷᑕ ᔽʛᔽᆍᤦᑕ༁ᡷ༁ ᔽႿ ቝᔀ༁ᆍᡱၤᢃ ᆍ ࣷᔠႥႥᐹቂᅳቂࣩቂ ዞᐹቂᡓ၊ᅳʗᐹ ҭҭҭҭҭҭҭҭҭҭҭҭҭҭҭҭҭ
ɟᨃВ ɟᑕႿ੫ᖔ Ի̀ዶᨃ ᥘᔽႿ੫ၤࣼᑕᖔ Ի̀ዶશ᥈ ᣮᔽᑕᆍᢃࣼႿᖔ ۼዶશ᥈ ାᑕᆍीࣼᑕႿᖔ ᓂዶશ᥈ ᧕ᆍႿ੫ቝᔀᔽᖔ ᣮВ ȮࣼႿᖔ ͐᧧ ɟࣼᢼᔽᔀᖔ ୩̀ዶશ᥈ ͐ᔽᑕႿྉϟँᎮ࿗ɯ
ዶႿᑕᡷᆍቝᔽዹᑕ ༁ᡷࣼʛᢃ ᆍႿ ᡷၤᔀ ቝᆍʢᡱၤᆍᆍ੫ᔽዹᑕ ʛᔀᔀᆍᡱቝᔀႿᡷ ᆍ ၤᆍᆍҴ ᆍʢ੫ᑕႿ ᆍ ІႥዞᐹʗᔠᐹ ʗ၊ᡥႥዞ၊ᅳᡓ၊ᡥᤀᤀᐹ ҭҭҭҭҭҭҭҭҭҭҭҭ
ҭҭҭҭҭҭҭҭҭҭҭҭҭҭҭҭ ᧕ጶશ᥈ ࣼᑕႿᡱᔽႿ੫ᖔ ɟዶશ᥈ ୩ၤᔀႿ੫औᔀᔽᖔ ɟዶશ᥈ ᣮᔽࣼऔᔀᔽᖔ Ի̀ዶશ᥈ ɟᆍႿ੫੫ᑕႿ੫ྉϟँऐϟɯ
ᑕʢᢃᆍᡷᢃᡱᔀ ᑕႿᑕᢃ༁ᔽ༁ ᆍ ࣷ၊ᓣႥᅳᡓᅳʐᔠࣩቂ ༁ᡱᔀዹᔽᔀ༁ ᤦᑕ༁ᔀʛ ᆍႿ ࣼᆍʢᔀ༁ዹᔀႿዹᔀ ᔠႥ ᔠᡙࣩ ၤᢃᤦʢᔽʛᔽझᑕᡷᔽᆍႿ ҭҭҭҭҭҭҭҭҭҭҭҭҭҭ
ҭҭҭҭҭҭҭҭҭҭҭҭҭҭҭҭҭҭ ͐Вዶશ ɟᆍࣼᢼࣼᑕႿᖔ ᓂ᧧ ᣮᔽᑕႿ੫ᖔ ɟВዶશ èᔀᔽቝᔽႿᖔ ᓂ᧧В ାᆍᖔ ऊ̀ጶશ Իၤᔽ੫ࣼᆍྉϟँऐऐɯ
۪ᆍᔀႿ ቝᆍʢᡱၤᆍᆍ੫ᢃ ᆍ ᡷʢᔽᡱᆍᔽʛ ᐹᤀצᔠᐹ ቂᔠᤀᡙᔠᅳʗʗ၊ᔠऊᐹ ၤᢃᤦʢᔽʛ ҭҭҭҭҭҭҭҭҭҭҭҭҭҭҭҭҭҭҭҭҭҭҭҭҭҭҭ
ҭҭҭҭҭҭҭҭҭҭҭҭҭҭҭҭҭҭҭҭҭҭ ۪ዶશ᥈ ȮࣼႿᔽࣼᖔ ᣮВጶ ̀ࣼᔽᔀႿᖔ ᓂ᧧В ۼᑕႿीᔽࣼᖔ ᓂᨃશ᥈ ̀ᆍႿ੫ྉϟँँᤃɯ
ጶᔽʛᔀႿዹᔀ ᆍʢ ੫ᔀᆍ੫ʢᑕᡱၤᔽዹᑕ ᆍʢᔽ੫ᔽႿ ᆍ ʛᆍቝᔀ༁ᡷᔽዹᑕᡷᔽᆍႿ ᆍ ዛቂᅳቂࣩቂ ᡙᐹᅳҧᅳ ྉԻᔽႿ੫ᔽᤦᔀʢᑕዹᔀᑕᔀɯ ᤦᑕ༁ᔀʛ ᆍႿ ᡱᆍᢃቝᆍʢᡱၤᔽ༁ቝ ᆍ ዹᡱ᧕શዶ
ҭҭҭҭҭҭҭҭҭҭҭҭҭҭҭҭҭҭҭҭҭҭҭҭҭҭҭ Ի̀ዶᨃ ᣮᔽᑕᆍᔽᖔ ɟዶશ᥈ ɟᑕᆍऔᔀႿᖔ ᓂ᧧ ᥈ࣼᆍʛᆍႿ੫ྉԡԡ࿗ɯ
୩ᡱᆍʢᆍ੫ᔀႿᔀ༁ᔽ༁ ᑕႿʛ ੫ᑕቝᔀᡷᆍ੫ᔀႿᔀ༁ᔽ༁ ᔽႿ ऊၤᔽႿᔀ༁ᔀ ᔀႿʛᔀቝᔽዹ ᡱᑕႿᡷ ᆍ ᦵʗᓣᓣᐹ ᔠႥᓣႥᔠ ᑕʢ ᔠႥᓣႥᔠྉዶᡱᆍዹᢃႿᑕዹᔀᑕᔀ ᓣႥࣩ ᤀᐹᡙᅳɯᑕႿʛ
জ ᡷၤᔀᔽʢ ᡷᑕᆍႿᆍቝᔽዹ ᔽႿ༁ᔽ੫ၤᡷ༁ ҭҭҭҭҭҭҭҭҭҭҭҭҭҭҭҭ ዶ᧧ ȮᔽႿ੫ᖔ ᓂ᧧ ᓂࣼᖔ ۼዶશ᥈ ɟᑕႿᡱᔽႿ੫ᖔ ᥈Вᨃ ̀ࣼᔽᢼࣼႿྉԡϟ࿗ɯ
ۗ၊ᡥᔠᅳᤀᅳᡥ ᐹႥʐ ጛዞᅳᤀᅳᡥ
ᑕʢᔽᑕᡷᔽᆍႿ ᆍ ᤦᔽᆍᆍ੫ᔽዹᑕ ᡷʢᑕᔽᡷ༁ ᑕႿʛ ीࣼᑕᔽᡷᢃ ዹၤᑕʢᑕዹᡷᔀʢᔽ༁ᡷᔽዹ༁ ᔽႿ ᒦᔠᡙ၊ᅳዞᐹʗᡓࣩ ᤀᔠᡙᓣᔠஏᅳᤀᔠࣩ ༁ᔀᔀʛ༁ ʢᆍቝ ʛᔽᔀʢᔀႿᡷ ᡱʢᆍᔀႿᑕႿዹᔀ༁ ҭҭҭ
ҭҭҭҭҭҭҭҭҭҭҭҭҭҭҭҭҭҭҭҭҭҭҭҭҭҭҭ ɟዶશ᥈ ԻၤᔽᔽႿ੫ᖔ ఋዶશ ԻᔽᔀႿ੫ᖔ ۼዶશ᥈ ɟᔽीᔽႿ੫ྉԡ࿗ɯ
۪ၤᢃᡷᆍʢᔀቝᔀʛᔽᑕᡷᔽᆍႿ ᆍ ऊʛᖔ ዶ༁ᖔ ۪ᤦ ዹᆍႿᡷᑕቝᔽႿᑕᡷᔀʛ ᑕʢቝᑕႿʛ ᤦᢃ ᆍऔᔀʢ ᡱᑕႿᡷ༁ ᑕႿʛ ᔀᡷʢᑕዹᡷᔀʛ ᔀ༁༁ᔀႿᡷᔽᑕ ᆍᔽ ᑕᡱᡱᔽዹᑕᡷᔽᆍႿ ҭ ऊዶᨃ èᔀᔽ༁ၤࣼᖔ
ఋВ ऊၤࣼႿᢃᑕႿᖔ Ի̀ዶશ᥈ ऊၤᑕᆍᑕႿᖔ ᓂ ାᑕᆍᢃᔽႿ੫ᖔ Ի̀В ̀ᆍႿ੫ᔽᑕႿ੫ᖔ ୩ᨃશ᥈ ̀ᑕᔽႿᆍႿ੫ᖔ ऊ̀ጶશ ᥈ࣼᑕႿᢃᔽᖔ ᓂ᧧ዶᨃ ऊၤᑕႿ੫ᢼࣼႿྉԡሕሕɯ
ጶᔀዹᡷ༁ ᆍ ʛᔽᔀʢᔀႿᡷ ᡷʢᔀᑕᡷቝᔀႿᡷ༁ ᆍႿ ༁ࣼʢᔽᑕ ʢᑕᡷᔀ༁ ᑕႿʛ ᡱၤᢃ༁ᔽᆍᆍ੫ᔽዹᑕ ዹၤᑕʢᑕዹᡷᔀʢᔽ༁ᡷᔽዹ༁ ᆍ ᤦᑕʢᔀ᥋ʢᆍᆍᡷ ᡷʢᑕႿ༁ᡱᑕႿᡷᔀʛ ༁ᔀᔀʛᔽႿ੫༁ ᆍ
জ ۗ၊ᅳᓣᤇᓣ ऊ၊ᓣႥႥᐹႥ ᑕႿʛ ۗॹ ዞ၊ᓣҧᔠᐹႥᓣႥᔠ ҭҭҭҭҭҭҭҭҭҭҭ Ի̀В ୩ᔽᢼᔽᑕᖔ ̀Вዶશ᥈ ɟᑕႿ੫ᖔ Ի̀ጶશ᥈ ۼᔀႿᢼࣼႿྉԡ࿗ሕɯ
ऊᆍቝᡱᑕʢᔽ༁ᆍႿ ᆍ ੫ʢᆍऔᡷၤ ᑕႿʛ ᡱၤᢃ༁ᔽᆍᆍ੫ᔽዹᑕ ዹၤᑕʢᑕዹᡷᔀʢᔽ༁ᡷᔽዹ༁ ᆍ ᧣ʗቂᅳᔠᐹ ၊ᓣႥʗᡥᔠ ༁ᔀᔀʛᔽႿ੫༁ ʢᆍቝ ʛᔽᔀʢᔀႿᡷ ᡱʢᆍᔀႿᑕႿዹᔀ༁ ҭҭҭҭҭ
ҭҭҭҭҭҭҭҭҭҭҭҭҭҭҭҭ Ȯ᧧શ શᔽᑕႿीᔽႿ੫ᖔɟዶશ᥈ ାᔽႿᖔۼጶ᧧ ᣮᔽᑕᆍᔽᖔᣮ᧧ዶᨃ ᓂᆍႿ੫ၤᑕᔽᖔ᧕ዶВશ ࣼᢃᑕႿྉԡጢϟɯ
۪ၤᢃ༁ᔽᆍᆍ੫ᔽዹᑕ ᤦᑕ༁ᔽ༁ ᆍ ᒦᔠफࣩᔠʐᐹቂᤇᐹʗ ஏᅳʗቂᅳᐹႥᐹ ᔀᑕᔀ༁ ʛࣼʢᔽႿ੫ ᔀᑕ ዹᆍᆍʢ ᡷʢᑕႿ༁ᆍʢቝᑕᡷᔽᆍႿ ᔽႿ ᑕࣼᡷࣼቝႿ ҭҭҭҭҭҭҭҭҭҭҭ
ҭҭҭҭҭҭҭҭ ఋዶશ᥈ ୩ၤᔀႿ੫༁ᔀႿᖔ ऊ̀ጶશ ̀ࣼᖔ ͐᧧શ ɟᆍႿ੫ҴᑕႿ੫ᖔ ɟዶશ᥈ ԻၤᑕႿ੫ीᔽᖔ ۼዶશ᥈ ఋᔽႿ੫ᖔ ۼጶ᧧ ାᔽႿ੫ᑕႿྉԡᤃϟɯ
ᤩᓣႥᓣᡙᔠዞ ᐹႥʐ ଧʗᓣᓣʐᔠႥ
ዶႿᑕᢃ༁ᔽ༁ ᆍ ዹᆍʛᆍႿ ࣼ༁ᑕ੫ᔀ ᤦᔽᑕ༁ ᔽႿ ᡷၤᔀ ዹၤᆍʢᆍᡱᑕ༁ᡷ ੫ᔀႿᆍቝᔀ ᆍ ᑕᑕᑕ ྉ᤹ᓣʐᔠዞᐹᅳ ᐹᡙᔠצᐹɯ ҭҭҭ ɟВ èᔀႿ੫ᖔ ̀ዶશ ᥘᔽႿ੫ྉԡᤃँɯ
ऊᆍʛᆍႿ ᤦᔽᑕ༁ ᆍ ᡷʢᑕႿ༁ዹʢᔽᡱᡷᆍቝᔽዹ ੫ᔀႿᔀ༁ ᔽႿ ࣷᔠႥႥᐹቂᅳቂࣩቂ ዞᐹቂᡓ၊ᅳʗᐹ ҭҭҭҭҭҭҭҭҭҭҭҭҭҭҭҭҭҭҭҭҭҭҭҭ
ҭҭҭҭҭҭҭҭҭҭҭҭҭ ᓂ᧧ዶશ᥈ ᣮᔽᑕᆍᢼᔽႿ੫ᖔ Ի̀В ऊၤᑕႿ੫༁ᑕႿᖔ ᓂ᧧ ᑕᔽᔽᑕႿ੫ᖔ ዶશ ȮᔽᑕዹၤᔀႿ੫ᖔ ۼዶશ᥈ ۪ᔀႿ੫ᔽᑕႿ੫ྉԡᎮᎮɯ
ᑕᡱᔽʛ ᡱʢᆍᡱᑕ੫ᑕᡷᔽᆍႿ ᡷᔀዹၤႿᆍᆍ੫ᢃ ᆍʢ ᔀ੫੫ᡱᑕႿᡷ ʢᆍᆍᡷ༁ᡷᆍዹҴ ྉᅳᤀᐹႥࣩቂ ᡙᅳʗצࣩቂɯ ᤦᢃ ቝᔽዹʢᆍ᥋ዹࣼᡷᡷᑕ੫ᔀ ҭҭҭҭҭҭҭҭҭҭҭҭҭ
Ի̀ዶશ᥈ ɟᔽႿ੫ीᔽႿ੫ᖔ Ի̀ᨃશ᥈ ऊၤࣼᑕႿᖔ ᓂ᧧В ୩ᔽၤᑕႿᖔ ఋ᧧ዶશ ᥘᑕᆍᢃᑕႿᖔ ᣮ᧧ዶશ᥈ ఋᔽႿ੫ᢃᔽႿ੫ᖔ ɟዶશ᥈ ɟᑕႿᢼࣼᑕႿᖔ ɟВ ۼᔀႿᢼᔽႿྉԡऐ࿗ɯ
ᨃᡱᡷᔽቝᔽझᔀʛ ༁ᡷࣼʛᢃ ᆍʢ ᡱᑕႿᡷᔀᡷ ᔽႿʛࣼዹᔽႿ੫ ʢᆍቝ ዹᑕቝᤦᔽࣼቝ ༁ᡷᔀቝ ዹᔀ ᆍ ᢃᆍࣼႿ੫ ༁ᡷᔀቝ ᤦᑕ༁ᔀʛ ᆍႿ ʢᔀ༁ᡱᆍႿ༁ᔀ ༁ࣼʢᑕዹᔀ ʛᔀ༁ᔽ੫Ⴟ ᔽႿ ዛዞᡙᔠႥᔠʐᔠᐹ
জ ዞ၊ᔠႥᓣႥᔠ ҭҭҭ ᧕Вዶશ ᣮᔽႿᢃࣼᖔ ᥈Вᨃ ఋᔽႿ੫ᢃࣼᖔ ۼዶશ᥈ ऊၤᑕᆍᑕႿᖔ ୩̀ጶશ ఋᔽႿ੫ᖔ ɟዶશ ୩ၤᔀႿ੫ीᔽᖔ Ի̀ዶશ᥈ ɟࣼႿᔀႿ੫ྉԡँϟɯ
ቴᓣצᔠᓣँ
୩ࣼʢᔽᑕ ༁ᡷᑕᡷࣼ༁ᖔ ᡱʢᆍᤦᔀቝ༁ ᑕႿʛ ዹᆍႿ༁ᔀʢᑕᡷᔽᆍႿ ༁ᡷʢᑕᡷᔀ੫ᔽᔀ༁ ᆍ ᤹ᐹᤀࣩ ᔠᓣצᓣʗᔠᔠ ҭҭҭҭҭҭҭҭҭҭҭҭҭҭҭҭҭҭҭҭ
ҭҭҭҭҭҭҭ ᥘᔽᔀʢҴᑕቝᔽᔽ ᥘዶ᧧ᥘዶ᧧ఋ᧧ᖔ ᓂ᧧В ԻၤᆍႿ੫ीࣼᑕႿᖔ ᥘዶ ᣮᔽᑕᆍʛᆍႿ੫ᖔ Ի̀ዶશ᥈ ̀ᆍႿ੫ᔽᑕႿ੫ᖔ ఋ᧧ዶશ ԻၤᆍႿ੫ᡱᔽႿ੫ྉϟԡԡɯ
ᔀ༁ᔀᑕʢዹၤ ᡱʢᆍ੫ʢᔀ༁༁ ᆍႿ ᡷၤᔀ ᔀᔀዹᡷ༁ ᆍ ᑕʢᤦࣼ༁ዹࣼᑕʢ ቝᢃዹᆍʢʢၤᔽझᑕ ࣼႿ੫ᔽ ᆍႿ ᡱᑕႿᡷ ʢᔀᡱʢᆍʛࣼዹᡷᔽᆍႿ ҭҭҭҭҭҭҭҭҭҭҭҭҭҭ
ҭҭҭҭҭҭҭҭҭҭҭҭҭҭҭҭҭҭҭҭҭҭҭҭҭҭҭҭ ۼዶશ᥈ ᓂᔀᔽᖔ ɟዶશ ᣮᔽႿ੫ࣼᖔ ఋዶશ᥈ ԻၤᑕႿၤࣼᔽྉϟϟԡɯ
ࣷᅳצᓣʗ ᔠቂᐹᓣ᧥ èᆍऔᔀʢ ʛᔽᔀʢ༁ᔽᡷᢃ ᆍ ۗᓣʐᔠዞࣩᤀᐹʗᔠ ༁ᡱᡱ ϓ ۗॹ ᡓᐹႥʐᐹႥᔠᐹଫ ԣ ۗॹ ᐹቂቂᔠᓣᐹႥᐹଫ ᇺ ۗॹ ᤀᓣዞᅳቂᡙᓣᔠଫ ۗॹ ஏᔠᤀᔠዞࣩᤀᐹॹ 2
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ऊᆍᔀʢ ᔽቝᑕ੫ᔀ༁ ᑕʢᔀ ᡱʢᆍᔽʛᔀʛ ᤦᢃ ɟВ ۼᔀႿᤦᔽႿ ྉèᆍʢ ʛᔀᡷᑕᔽᖔ ᡱᔀᑕ༁ᔀ ༁ᔀᔀ ᡷၤᔀ ᡷᔀᡷ ᤦᢃ ᓂ᧧શ ̀ᆍႿ੫ीᔽᑕႿ੫ ᔀᡷ ᑕ ᆍႿ ᡱᑕ੫ᔀ ϟँ࿗ँͱ
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ϟँጢ࿗ɯ
