Page 4 - 《广西植物）》2021年第1期

ইᤩІᧇ̵ዛᧇዛইইইইইইইইইইই ȣࣩႥॹ ԣԔԣԔই Ⴌᅳᤀᤫ ྽ϓ ટᅳᤫ ϓ

ইইইইইইইইইইইইইইইইইইইইই ྉ᤹ᅳႥᡙ၊ᤀᡥɯ

ࣷ᧣ટ௴ጛટ௴୒

୒ᡓᓣዞᔠᐹᤀ ᧇ໧໧ࣩᓣ᧥ ۗᤀᐹႥᡙ ዞᤀᐹ໧໧ᔠஏᔠዞᐹᡙᔠᅳႥ ᐹႥʐ ୒ᡥ໧ᡙᓣቂ ጛצᅳᤀࣩᡙᔠᅳႥ

᧕ᔽ༁ዹᆍ׹ᔀʢᢃ ᆍ஦ ࣷᤀᓣቂᐹᡙᔠ໧ ᡙᅳႥ੖ᤀࣩᓣႥ໧ᔠ໧ ྉ኏ᑕႿࣼႿዹࣼ᤟ᑕዹᔀᑕᔀɯ ஦ʢᆍቝ ୩ᆍࣼᡷၤᔀᑕ༁ᡷ ఋᔽᤦᔀᡷ ᆍ஦ ऊၤᔽႿᑕ ҭҭҭҭҭҭҭ ۼዶશ᥈ ۼᔀႿ᥋ᡷ༁ᑕᔽྉϟɯ
௴ᓣࣩዞʗᔠࣩቂ फᔠႥ੖ᡥࣩᐹႥᓣႥ໧ᓣᖔ ᑕ Ⴟᔀऔ ༁ᡱᔀዹᔽᔀ༁ ᆍ஦ ᓂᑕቝᔽᑕዹᔀᑕᔀ ஦ʢᆍቝ ԻၤᔀᢼᔽᑕႿ੫ᖔ ऊၤᔽႿᑕ ҭҭҭҭҭҭҭҭҭҭҭҭҭҭҭҭҭҭҭ
ҭҭҭҭҭҭҭҭҭҭҭҭҭҭҭҭҭҭҭҭҭҭ ऊ̀ጶશ ᧕ᔀ᤟ᔽᑕႿ੫ᖔ Ȯ᧧શ ᣮᔽᑕᆍ஦ᔀႿ੫ᖔ ᣮВ ɟࣼᔀ᤟ᔽᑕႿ੫ᖔ ᧕᧧શ᥈ ାᔽႿ੫ᢃᑕႿ੫ྉ࿗ɯ
᧣໧ቂᐹႥᡙ၊ࣩ໧ ᐹࣩ໧ᡙʗᅳऊ၊ᓣᢝᔠᐹႥ੖ᓣႥ໧ᔠ໧ ྉᨃ᤟ᔀᑕዹᔀᑕᔀɯᖔ ᑕ Ⴟᔀऔ ༁ᡱᔀዹᔽᔀ༁ ஦ʢᆍቝ ԻၤᔀᢼᔽᑕႿ੫ ҭҭҭҭҭҭҭҭҭҭҭҭҭҭҭҭҭҭҭ
ҭҭҭҭҭҭҭҭҭҭҭҭҭҭҭҭҭҭ ᣮ᧧ጶ ۼᔀႿᢃࣼᑕႿᖔ ᓂ᧧В ᣮᔽᖔ ᥘጶ᧧ ᣮࣼʛᆍႿ੫ᖔ ऊ̀ጶશ èᔀႿ੫ᖔ ऊ̀ጶશ ԻၤᔀႿ੫ၤᑕᔽྉϟԡɯ
ዶ Ⴟᔀऔ ༁ᡱᔀዹᔽᔀ༁ ᑕႿʛ ᑕ Ⴟᔀऔ ʢᔀዹᆍʢʛᔀʛ ༁ᡱᔀዹᔽᔀ༁ ᆍ஦ ̵ᐹᓣቂᐹᡙᅳቂቂᐹ ஦ʢᆍቝ ऊၤᔽႿᑕ ྉᓂᔀዹᑕႿᆍʢᑕ᤟ᔀ༁᧥ ̀ᑕᔀቝᑕᡷᆍቝቝᑕᡷᑕዹᔀᑕᔀɯ ҭҭҭҭҭҭ
ҭҭҭҭҭҭҭҭҭҭҭҭҭҭҭҭҭҭҭҭҭҭҭ ኏ጶશ Իၤᑕᆍᢼᔽᔀᖔ ఋዶશ᥈ ኏ᆍႿ੫ᖔ ᧕ᨃશ᥈ ᓂᔽႿ᤟ᔽႿᖔ Ի̀ዶશ᥈ ᓂࣼ᤟ࣼྉϟᤃɯ
ዶʛ׹ᑕႿዹᔀ༁ ᔽႿ ᡷၤᔀ ༁ᡷࣼʛᢃ ᆍ஦ ༁ᢃ༁ᡷᔀቝᑕᡷᔽዹ༁ ᆍ஦ ᥥጶᑕ༁ᡷ᥋ዶ༁ᔽᑕ ऊ᤟ᑕʛᔀɼ ᔽႿ ዶᡱᔽᑕዹᔀᑕᔀ ҭҭҭҭҭҭҭҭҭҭҭҭҭҭҭҭҭҭҭҭ
ҭҭҭҭҭҭҭҭҭҭҭҭҭҭҭҭҭҭҭҭҭҭҭҭҭҭҭҭҭ Ի̀ᨃВ ȮᔽႿ੫ᖔ ᥈Вᨃ ᥘᔽႿ੫ᢼᔽᑕᖔ ᓂ᧧В ԻၤᔀႿऔᔀႿྉ԰ϟɯ
ዶʛ׹ᑕႿዹᔀ༁ ᔽႿ ቝᆍ᤟ᔀዹࣼ᤟ᑕʢ ᡱၤᢃ᤟ᆍ੫ᔀႿᢃ ᆍ஦ ᧇʗᔠ໧ ҭҭҭҭҭҭҭҭҭҭҭҭҭҭҭҭҭҭҭҭҭҭҭҭҭҭҭҭҭҭҭҭҭ
ҭҭҭҭҭҭҭҭ ऊ̀ጶશ᥈ᓂᔽႿᖔ èጶશ᥈୩ၤࣼዹၤᔀႿ੫ᖔ ᣮ᧧ዶᨃɟࣼᔀᆥᔀᖔ ɟВèᔀႿ੫ᢃᑕႿ੫ᖔ Ի̀В୩ၤࣼ଴ᔽᑕᖔ ۼዶશ᥈ᣮᔽᑕႿʢᆍႿ੫ྉሕϟɯ
᥈ᔀႿᆍቝᔀ᥋औᔽʛᔀ ᔽʛᔀႿᡷᔽ஦ᔽዹᑕᡷᔽᆍႿ ᑕႿʛ ᡱၤᢃ᤟ᆍ੫ᔀႿᔀᡷᔽዹ ᑕႿᑕ᤟ᢃ༁ᔽ༁ ᆍ஦ ୩ۼጶጶఋ ᡱʢᆍᡷᔀᔽႿ ஦ᑕቝᔽ᤟ᢃ ᔽႿ ᡱࣼቝᡱҴᔽႿ ҭҭҭҭҭҭҭҭҭҭҭҭ
ҭҭҭҭҭҭҭҭҭҭҭҭҭҭҭҭҭҭҭҭҭҭҭҭҭҭҭҭҭҭҭҭ ୩̀ጶશ ऊၤᑕႿ੫औᔀᔽᖔ ɟВዶશ ȮᔽႿ੫ᡱᔽႿ੫ྉ࿗ԡɯ
ዶ ዹ᤟ࣼ༁ᡷᔀʢ ᑕႿᑕ᤟ᢃ༁ᔽ༁ ᆍ஦ Ҵᑕʢᢃᆍᡷᢃᡱᔀ ʢᔀ༁ᔀቝᤦ᤟ᑕႿዹᔀ᥋Ⴟᔀᑕʢ ዹᆍᔀ஦஦ᔽዹᔽᔀႿᡷ༁ ᔽႿ ੫ᔀႿࣼ༁ ጛᡓᔠቂᓣʐᔠࣩቂ ᓂ঎ ᑕႿʛ ᔽᡷ༁ ᔀ׹ᆍ᤟ࣼᡷᔽᆍႿᑕʢᢃ ᑕႿʛ ༁ᢃ༁ᡷᔀቝᑕᡷᔽዹ
জ ᔽቝᡱ᤟ᔽዹᑕᡷᔽᆍႿ༁ ҭҭҭҭҭҭҭҭҭҭҭҭҭҭҭҭҭҭҭ ऊ̀ጶશ ɟᆍႿ੫ᤦᔽᖔ ᓂ᧧ ୩ၤࣼᑕႿ੫ᖔ ۼВ ȮᔽႿ੫ᖔ ୩̀ጶશ᥈ ᥘᑕᆍᢃᔽႿྉጢጢɯ
۪ၤᢃ᤟ᆍ੫ᔀႿᢃ ᆍ஦ ᥘᢃʢᡷᑕ᤟ᔀ༁ ᑕႿʛ ʢᔀ᤟ᑕᡷᔀʛ ੫ʢᆍࣼᡱ༁ ᤦᑕ༁ᔀʛ ᆍႿ ዹၤ᤟ᆍʢᆍᡱ᤟ᑕ༁ᡷ ੫ᔀႿᆍቝᔀ ҭҭҭҭҭҭҭҭҭҭҭҭҭҭҭҭҭҭҭҭ
ҭҭҭҭҭҭҭҭҭҭҭҭҭҭҭ ۼዶશ᥈ ᣮࣼᔀीᔽႿᖔ ୩ᨃશ᥈ ۼᔀᔽऔࣼᖔ ᣮ᧧ዶᨃ ȮᔽᑕႿᢼᔽᑕᖔ ᓂ᧧ ऊၤᑕᆍीᔽᆍႿ੫ᖔ ᓂ᧧В ̀ᆍႿ੫झၤᑕႿྉᤃऐɯ
ዶ ੫ᆍᆍʛ ቝᑕᡷᔀʢᔽᑕ᤟ ஦ᆍʢ ዹၤʢᆍቝᆍ༁ᆍቝᔀ ዹᆍࣼႿᡷ༁ ᆍ஦ ஦ᔀʢႿ༁ ҭҭҭҭҭҭҭҭ ۼዶશ᥈ ԻᔽᢼࣼᑕႿᖔ ᣮ᧧ዶશ᥈ ȮᔽᑕႿᢃᔽႿ੫ᖔऊ̀ጶશ᥈ ᣮᔽᑕᆍྉऐϟɯ
ऊၤᑕʢᑕዹᡷᔀʢᔽ༁ᡷᔽዹ༁ ᆍ஦ ༁ᡱᆍʢᔀ ੫ᔀʢቝᔽႿᑕᡷᔽᆍႿ ᑕႿʛ ᡱʢᆍᡷᆍႿᔀቝᑕ ʛᔀ׹ᔀ᤟ᆍᡱቝᔀႿᡷ ᆍ஦ ᡷऔᆍ ༁ᡱᔀዹᔽᔀ༁ ᔽႿ ÞᐹᤇʗᅳႥᔠᐹ ኏ᑕʛʛᔽ ҭҭҭҭҭҭҭҭҭ
ҭҭҭҭҭҭҭҭҭ ̀Вዶશ᥈ ୩ၤᔽ᤟ᔽᑕႿ੫ᖔ ऊዶᨃ ԻၤᔀႿᖔ Ի̀ዶશ᥈ ̀ᑕᆍᖔ શ᧧В ɟࣼ᤟ࣼᖔ ۼዶશ᥈ ԻၤᔀႿᢼᔽᔀᖔ Ի̀ዶᨃ ȮᔽᑕႿዹၤᔀႿ੫ྉऐጢɯ
ዶ Ⴟࣼቝᔀʢᔽዹᑕ᤟ ᡷᑕ଴ᆍႿᆍቝᔽዹ ᑕႿᑕ᤟ᢃ༁ᔽ༁ ᆍ஦ ᤩᤀᡥᡓᡙᅳᡓᓣᡙᐹᤀࣩቂ ᑕႿʛ ጛࣩᅳႥᡥቂࣩ໧ ྉऊᔀ᤟ᑕ༁ᡷʢᑕዹᔀᑕᔀɯ ҭҭҭҭҭҭҭҭҭҭҭҭҭҭҭҭҭ
ҭҭҭҭҭҭҭҭҭҭҭҭҭҭҭҭҭҭҭҭҭҭҭҭҭҭҭҭ ᓂ᧧В ̀ࣼᔽᢃࣼᑕႿᖔ ᥘጶશ᥈ ୩ၤᔽᢃᆍႿ੫ᖔ ᓂ᧧В ͐ࣼᑕႿʢࣼྉँ԰ɯ
۪ᆍ᤟᤟ᔀႿ ቝᆍʢᡱၤᆍ᤟ᆍ੫ᢃ ᑕႿʛ Ⴟࣼቝᔀʢᔽዹᑕ᤟ ᡷᑕ଴ᆍႿᆍቝᢃ ᆍ஦ ԰԰ ̵ᔠᤇᔠ໧ዞࣩ໧ ໧ᡥʗᔠᐹዞࣩ໧ ҭҭҭҭҭҭҭҭҭҭҭҭҭҭҭҭҭҭҭҭҭҭ
ҭҭҭҭҭҭҭ Ի̀ዶᨃ ɟᔽ଴ࣼᑕႿᖔ èጶશ᥈ ͐ᔽᖔ ఋ᧧ዶશ ᓂᔽႿᖔ Ի̀ዶશ᥈ Ȯᔽᑕᡱᔀᔽᖔ ۼዶશ᥈ ᣮᔽႿᖔ ᓂ᧧В ɟᔽዹၤᑕᆍᖔ ᓂ᧧В ᧕ᆍႿ੫ᢃࣼႿྉϟԡሕɯ
۪ၤᢃ᤟ᆍ੫ᔀႿᔀᡷᔽዹ ᡱᆍ༁ᔽᡷᔽᆍႿ ᆍ஦ ۗᅳႥዞᔠʗࣩ໧ ᑕႿʛ ۗᅳႥዞᔠʗࣩ໧ ᡓᅳᤀᡥᐹႥʐʗᐹ ᤦᢃ ᧕શዶ ༁ᔀीࣼᔀႿዹᔽႿ੫ ҭҭҭҭҭҭҭҭҭҭҭҭҭҭҭҭҭ
ҭҭҭҭҭҭҭҭҭҭҭҭҭҭҭҭҭҭҭҭҭҭҭҭҭҭҭҭҭ Ի̀ዶશ᥈ ɟࣼᖔ ᥈ᨃશ᥈ ᣮࣼႿᖔ èጶશ᥈ ᣮᔽࣼᢃᑕႿྉϟϟ࿗ɯ
ऊᆍቝᡱᑕʢᑕᡷᔽ׹ᔀ ᔽႿ׹ᔀ༁ᡷᔽ੫ᑕᡷᔽᆍႿ ᆍ஦ ׹ᔀ༁༁ᔀ᤟ ᡱᔀʢ஦ᆍʢᑕᡷᔽᆍႿ ᡱ᤟ᑕᡷᔀ༁ ᆍ஦ ኏ᑕႿࣼႿዹࣼ᤟ᔀᑕᔀ ྉ኏ᑕႿࣼႿዹࣼ᤟ᑕዹᔀᑕᔀɯ ᤦᑕ༁ᔀʛ ᆍႿ ʛᔽ஦஦ᔀʢᔀႿᡷ ቝᔀᡷၤᆍʛ༁ ҭҭҭ
ҭҭҭҭҭҭҭҭҭҭҭҭҭҭҭҭҭҭҭҭҭҭҭҭҭҭҭҭҭҭҭҭҭҭҭҭҭҭҭҭ ᓂВᨃ ᥘᔽႿʢᆍႿ੫ྉϟ԰ሕɯ
В᤟ᡷʢᑕ༁ᡷʢࣼዹᡷࣼʢᑕ᤟ ዹၤᑕʢᑕዹᡷᔀʢᔽ༁ᡷᔽዹ༁ ᆍ஦ Ԯᔠऊᔠᡓ၊ࣩ໧ ᢝࣩᢝࣩᤇᐹ ዹ׹঎ ᓂᔽႿ੫औࣼዹၤᑕႿ੫झᑕᆍ ஦ʢࣼᔽᡷ ׹ᑕ༁ዹࣼ᤟ᑕʢ ᤦࣼႿʛ᤟ᔀ༁ ᡱၤ᤟ᆍᔀቝ ᑕႿʛ ᔽᡷ༁ ༁ࣼʢʢᆍࣼႿʛᔽႿ੫
জ ᡱᑕʢᔀႿዹၤᢃቝᑕ ዹᔀ᤟᤟༁ ҭҭ ̀Вዶશ᥈ ɟࣼᔀᖔ Ի̀ዶશ᥈ ɟᔽႿ੫ዹᑕᔽᖔ ୩В ۼᔀᔽʛᆍႿ੫ᖔ ̀ዶ᧧ ɟࣼᑕႿᖔ Ի̀ዶશ᥈ ɟࣼᑕႿᖔ Ȯ᧧શ᥈ ̀ᆍႿ੫଴ᔽᑕྉϟሕሕɯ
ዶႿᑕᡷᆍቝᢃ ᑕႿʛ ၤᔽ༁ᡷᆍዹၤᔀቝᔽዹᑕ᤟ ༁ᡷᑕᔽႿᔽႿ੫ ᆍ஦ ᑕ᤟᤟ᔽ੫ᑕᡷᆍʢ औᔀᔀʛ ྉዛᤀᡙᓣʗႥᐹႥᡙ၊ᓣʗᐹ ᡓ၊ᔠᤀᅳଝᓣʗᅳᔠʐᓣ໧ɯ ҭҭҭҭҭҭҭҭҭҭҭҭҭҭҭҭ
ҭҭҭҭҭҭҭҭ ۼዶશ᥈ ఋᔽႿ੫ᖔ Ի̀ዶશ᥈ ᣮᔽᑕᖔ Ի̀ᨃВ ऊࣼႿᢃࣼᖔ ̀В ୩ၤࣼᑕႿ੫༁ၤࣼᑕႿ੫ᖔ ఋዶશ᥈ ɟࣼᖔ ɟዶશ᥈ ऊၤᑕᆍʛᆍႿ੫ྉϟ࿗࿗ɯ
ᥘᔽዹʢᆍ༁ዹᆍᡱᔽዹ ዹၤᑕʢᑕዹᡷᔀʢᔽ༁ᡷᔽዹ༁ ᆍ஦ ᤟ᔀႿᡷᔽዹᔀ᤟ᖔ ༁ᡷᆍቝᑕᡷᑕ᤟ ᑕᡱᡱᑕʢᑕᡷࣼ༁ ᑕႿʛ ᤟ᆍऔᔀʢ ᔀᡱᔽʛᔀʢቝᔽ༁ ᆍ஦ ᤟ᔀᑕ஦ ஦ʢᆍቝ ஦ᔽ׹ᔀ Ҵᔽऔᔽ஦ʢࣼᔽᡷ ༁ᡱᔀዹᔽᔀ༁ ҭҭҭҭ
ҭҭҭҭҭҭҭҭҭҭҭ ᓂ᧧В ۪ᔽႿ੫ᡱᔽႿ੫ᖔ ᣮ᧧ᨃશ᥈ ɟᑕ᤟ᑕႿᖔ ۼጶ᧧ ɟࣼᢼᔽႿ੫ᖔ ᥘᨃ ͐ࣼᑕႿၤࣼᔽᖔ ۼዶશ᥈ èᑕቝᔽႿ੫ᖔ ɟጶ ᣽ᑕᔽᢃࣼྉϟጢᎮɯ

ࣷᅳצᓣʗ ᔠቂᐹ੖ᓣ໧᧥ ୩ᔀ᤟ᔀዹᡷᔽ׹ᔀ ᡱʢᔀ༁ᔀႿᡷᑕᡷᔽᆍႿ ᆍ஦ ऊᔀ᤟ᑕ༁ᡷʢᑕዹᔀᑕᔀ ᡱ᤟ᑕႿᡷ༁঎ ϟ঎ ጛࣩᅳႥᡥቂࣩ໧ ᅳଝᡥᡓ၊ᡥᤀᤀࣩ໧ଫ԰঎ ጛॹ ၊ᐹቂᔠᤀᡙᅳႥᔠᐹႥࣩ໧ଫሕ঎ ጛॹ 1 2
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ஏʗᔠ੖ᔠʐࣩ໧ ׹ᑕʢ঎ ዞᅳʗႥࣩᡙᅳᔠʐᓣ໧ଫ࿗঎ ጛॹ ዞᐹʗႥᅳ໧ࣩ໧ଫጢ঎ ጛॹ ᤀᐹଝᔠஏᤀᅳʗࣩ໧঎ ऊᆍ׹ᔀʢ ᔽቝᑕ੫ᔀ༁ ᑕʢᔀ ᡱʢᆍ׹ᔽʛᔀʛ ᤦᢃ Իၤࣼ ᣮᔽႿ଴ᔽႿ঎ྉèᆍʢ ʛᔀᡷᑕᔽ᤟ᖔ 4
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ᡱ᤟ᔀᑕ༁ᔀ ༁ᔀᔀ ᡷၤᔀ ᡷᔀ଴ᡷ ᤦᢃ ᓂ᧧В ̀ࣼᔽᢃࣼᑕႿ ᔀᡷ ᑕ᤟঎ ᆍႿ ᡱᑕ੫ᔀ ँ԰ͱϟԡ԰ɯ