中国花椒属(广义)叶结构研究

曹 明<sup>1; 2; 3</sup>; 曹丽敏<sup>4</sup>; 张奠湘<sup>2*</sup>; 张瑞泉<sup>1</sup>

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中国花椒属(广义)叶结构研究
曹明^1,2,3, 曹丽敏⁴, 张奠湘^2*, 张瑞泉¹
1. 广西壮族自治区中国科学院广西植物研究所, 广西桂林 541006;2.中国科学院华南植物园, 广州 510650;3.中国科学院研究生院, 北京 100049;4.湖南衡阳师范学院生命科学系, 湖南衡阳 421008

摘要:

对国产花椒属32种4变种叶结构特征进行了研究。结果发现,花椒属叶脉均属于羽状脉弓形脉序类型; 叶齿具全缘、具齿或钝齿3种类型; 二级叶脉有分支或不分支; 三级叶脉有分支、网状和及顶3种类型; 多数种类叶齿无腺点,仅4种叶齿有腺点。叶结构研究结果支持花椒亚属和崖椒亚属的亚属等级,它们共同组成广义的花椒属,并支持花椒属为芸香科中比较原始类群的观点; 但不支持Engler将花椒亚属和崖椒亚属独立成属的分类处理。基于重要外部形态学特征和叶结构特征,重新对花椒属下两个亚属的检索表进行了修订。

关键词: 芸香科花椒属叶结构中国

DOI：

分类号:Q944

Fund project:

Leaf architecture of Zanthoxylum(Rutaceae)fromChina and its taxonomic significance

CAO Ming^1,2,3, CAO Li-Min⁴, ZHANG Dian-Xiang^2*, ZHANG Rui-Quan¹

1.Guangxi Institute of Botany, Guangxi Zhuang Autonomous Region and the Chinese Academy of Sciences, Guilin 541006, China;2.South China Botanical Garden, the Chinese Academy of Sciences, Guangzhou 510650, China;3.Graduate School of the Chinese Academy of Sciences, Beijing 100049, China;4.Department of Life Sciences, Hengyang Normal University, Hengyang 421008, China

Abstract:

Leaf architecture of 32 species and 4 variety of Zanthoxylum s.l.(Rutaceae)in China were investigated. A comprehensive description of leaf architecture for Zanthoxylum was provided. Venation pattern was brochidodromous which represented a comparative primitive type. Tree types of teeth were detected,namely entire,toothed and crenate. The secondary veins were branched or unbranched. The most species were without glands on teeth,except in 4 species. These results did not support Englerian classification of Zanthoxylum. Based on comparison of leaf architecture,our results supported the taxonomic treatment of subgenera Fagara and Zanthoxylum as subgenus rank of Zanthoxylum and the genus Zanthoxylum as a comparative primitive group within the Rutaceae. A key based on main diagnostic morphologically and leaf architecture characters to the two subgenera of Zanthoxylum were revised.

Key words: Rutaceae Zanthoxylum leaf architecture China