| 摘要: |
| 首次报道了湖北双蝴蝶小孢子发生和雄配子体发育。主要结果如下：花药四室；药壁发育为双子叶型；绒毡层异型起源，属腺质型绒毡层，药隔处的绒毡层细胞形成类胎座，其余部位的绒毡层细胞仍为一层细胞；花药成熟时，药室内壁纤维状加厚且柱状伸长，表皮细胞减缩退化，纤维状加厚不明显。小孢子母细胞减数分裂为同时型，四分体排列方式主要为四面体形，少数为十字交叉形；成熟花粉多为2-细胞型，偶见3-细胞型，具三萌发孔。 |
| 关键词: 湖北双蝴蝶 小孢子 雄配子体 龙胆科 |
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| Genesis of microspore and the development of male gametophyte in Tripterospermum discoideum |
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HUANG Heng-Yu， LONG Hua， Yi Ting-Ting， LI Li*
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Key Laboratory of Plant Resources Conservation and Utilization（Jishou University）， College of Hunan Province， Jishou 416000， China
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| Abstract: |
| The present paper firstly reports the microsporogenesis and the development of male gametophyte in Tripterospermum discoideum. The main results can be concluded as follows:Anthers are tetrasporangiate. The development of anther walls conforms to the Dicotyledonous type and comprises of epidermis，endothecium，one or two middle layers and tapetum at the mature stage. The tapetal cells have dual origin and belong to the glandular type. The tapetal cells on the connective side show radial elongation or periclinal division and intrude into the anther locule to form placenoids；The endothecium persists and its cells become pillar and fibrous，and the epidermis degenerates. Cytokinesis at meiosis of microsporocytes is of the simultaneous type and most of microspore tetrads are tetrahedral，there are still a few other types，such as dilateral. Pollen grain is mainly 2 celled type when shed，occasionally 3 celled type，and has three apertures. |
| Key words: Tripterospermum discoideum microspore male gametophyte Gentianaceae |
