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8 广 西 植 物 40 卷
Basionym: Trapa japonica Flerov var. magnico ̄ Skvortzovꎬ in Bull. Jard. Bot. Princip. 26: 630. 1927. ≡
rona Z. T. Xiongꎬ in J. Wuhan Bot. Res. 3: 161. T. manshurica Flerov f. komarovii (Skvortzov) S.H. Li &
1985. Type: CHINAꎬ Hubeiꎬ Yangxinꎬ 1982ꎬ Y.L. Changꎬ in Fl. Pl. Herb. Chin. Bor. Or. 6: 137.
Z. T. Xiong 316 (holotype: WH!). 1977. Type: CHINAꎬ near Harbin and Taolaizao Stationꎬ
Synonym: Trapa japonica Flerov var. collector and number unknown.
longicollum Z. T. Xiongꎬ in J. Wuhan Bot. Res. 3: Synonym: Trapa acornis Nakanoꎬ in Bot.
161. 1985. Type: CHINAꎬ Hubeiꎬ Xiaoganꎬ 1982ꎬ Mag. (Tokyo) 77: 165. 1964. ≡ T. bicornis Osbeck
Z. T. Xiong 443 (holotype: WH!)ꎻ syn. nov. var. acornis ( Nakano) Z. T. Xiongꎬ in J. Wuhan
Trapa litwinowii V. N. Vassil. in Komarovꎬ Bot. Res. 3: 160. 1985. Type: CHINAꎬ near Shang ̄
Fl. URSS 15: 694. 1949. Type: RUSSIAꎬ Vallis haiꎬ Nakano s. n. ( holotype: Nakano Herbarium)ꎻ
fl. Ussuri inter Dsoadsa et Kindaꎬ 18 Aug. 1855ꎬ syn. nov.
C. Maximowicz s.n. (holotype: LE!). Trapa quadrispinosa Roxb.ꎬ Hort. Bengal.
Trapa litwinowii V. N. Vassil. var. chihuensis 11. 1814. ≡ T. bicornis Osbeck var. quadrispinosa
S. F. Guan et Q. Langꎬ Bull. Bot. Res. ( Harbin) 7 ( Roxb.) Z. T. Xiongꎬ in J. Wuhan Bot. Res. 3:
( 1 ): 77. 1987. Type: CHINAꎬ Jiangxiꎬ Chihu 160. 1985. Type: KASHMIRꎬ Dal Lakeꎬ 2 Aug.
Lakeꎬ 20 Sept. 1984ꎬ Y. D. Chenꎬ S. F. Guan & 1917ꎬ R. R. Stewart 3315 / 2 ( lectotype: Kꎬ desig ̄
Q. Lang 779 (holotype: PE!). nated by Vassiljev in 1960)ꎻ syn. nov.
冠菱 Fig. 3: C Trapa quadrispinosa Roxb. var. yongxiuensis
This variety is similar to Trapa natans var. natans W. H. Wanꎬ in J. Jiangxi Univ. (Nat. Sci.) 15(2):
in having the fruits with conspicuous beaksꎬ crowns 75. 1991. Type: CHINAꎬ Jiangxiꎬ Yongxiuꎬ
large and reflexedꎬ but differs in having the fruits lat ̄ Jiangyiꎬ W. H. Wan 780008 ( holotype: JXU!)ꎻ
erally compressedꎬ with two lower horns degenerated. syn. nov.
Additional collections studied. China. Hei ̄ 四角菱 Fig. 3:Dꎬ H
longjiang: Harbinꎬ C. Z. Zheng 6332 ( HZU )ꎬ This variety has 4 ̄horned fruitsꎬ but differs from
Y. L. Chang & S. D. Zhao 2766 ( IFP)ꎻ Qiqihaerꎬ Trapa natans var. natans in having fruits inconspi ̄
Z. S. Qin & C. F. Fang 101 ( IFP )ꎻ Songjiangꎬ cuously beaked and crowned. The fruit bodyꎬ horn
D. C. Zhao et al. 1310 ( IFP )ꎬ Y. L. Chou 1310 shape and lengthꎬ and exocarp color of T. natans
( IBSC ). Hubei: Wuchangꎬ Tangsun Lakeꎬ B. Y. var. komarovii are various and unstableꎬ especially in
Ding & R. Y. Hu 6261 ( HZU). Jiangxi: Yongxiuꎬ cultivated individuals. This variety is wildly distributed
W. H. Wan s.n. (JXU). Jilin: Fuyuꎬ S. D. Zhao & in Chinaꎬ as well as Indiaꎬ and many cultivated types
Y. L. Chang 2734 (IFP)ꎬ 2742 (IFP!)ꎬ Y. L. Chang occur now.
& S. D. Zhao 2758 (IFP)ꎬ 2759 (IFP)ꎬ S. D. Zhao Additional collections studied. China. Anhui:
& Y. L. Chang 2735 (IFP). Quanjiaoꎬ East China Exped. 3506 ( PE)ꎻ Wuhuꎬ
Distribution. China (Heilongjiangꎬ Henanꎬ Hu ̄ B. Y. Ding et al. 6217 (HZU). Fujian: without pre ̄
beiꎬ Jiangxiꎬ Jilinꎬ Liaoning )ꎬ Japanꎬ Korea and cise localityꎬ S. T. Dunn 2726 ( IBSC). Hainan:
Russia. Lingshuiꎬ Z. S. Diao s. n. ( YZU). Hebei: Anxinꎬ
Note. Trapa japonica var. magnicoronaꎬ T. Baiyangdianꎬ B. Y. Ding 6340 (HZU). Hubei: Jing ̄
japonica var. longicollumꎬ T. litwinowii var. chihuensis menꎬ H. Migo s. n. ( IBSC )ꎻ Wuchangꎬ Tangsun
and T. macropoda var. bispinosaꎬ which had the fruits Lakeꎬ B. Y. Ding & R. Y. Hu 6246 ( HZU ).
with conspicuous beaks and crowns ( Xiongꎬ 1985ꎻ Hunan: Jiangpuꎬ Z. S. Diao 2022 ( YZU ).
Guan & Langꎬ 1987ꎻ Wanꎬ 1991). These four varieties Jiangsu: Hongzeꎬ B. Y. Ding & X. D. Lu 6440 (HZU)ꎬ
are synonymized hereꎬ as well as T. octotuberculata. B. Y. Ding et al. 6232 (HZU)ꎻ Jiangningꎬ anonymous
2c. Trapa natans L. var. komarovii (Skvortzov) 6591 (NAS)ꎻ Nanjingꎬ J. J. Gong 801 (NAS)ꎻ Wuxiꎬ
B. Y. Ding & X. F. Jinꎬ comb. nov. Y. W. Law 3029 ( NAS ). Jiangxi: Nanchangꎬ W.
Basionym:Trapa amurensis Flerov var. komarovii H. Wan 79006 (JXU)ꎻ Yongxiuꎬ W. H. Wan 780008