Page 16 - 《广西植物》2020年第1期
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1 2                                    广  西  植  物                                         40 卷
       This variety is both in wild water or cultivation. Two  K. S. Hao 3271 ( PE )ꎬ 3288 ( PE )ꎻ Mt.
   cultivated groups were recognized:Trapa bicornis group  Funiushanꎬ Henan For. Bur. Exped. 285 ( PE ).
   and T. bispinosa group.                           Hubei: Shashiꎬ B. Y. Ding et al. 6250 ( HZU)ꎻ
       2e. Trapa natans L.     var.  quadricaudata   Wuchangꎬ Tangsun Lakeꎬ B. Y. Ding & R. Y. Hu
   (Glück.) B. Y. Ding & X. F. Jinꎬ comb. nov.       6263 ( HZU). Hunan: Hanshouꎬ W. X. Wang 14

       Basionym: Trapa incisa Siebold & Zucc. var.   (WH)ꎻ Hengyangꎬ B. Y. Ding & T. Huang 6390
   quadricaudata Glück in Handel ̄Mazzattiꎬ Symb.     (HZU)ꎻ Xiangyinꎬ B. Y. Ding & R. Y. Hu 6266
   Sin. 7 ( 2): 605. 1929. Type: CHINAꎬ Sichuanꎬ     (HZU). Jiangsu: Baoyingꎬ Baima Lakeꎬ S. L. Liu et
   Ningyüenꎬ 27 Oct. 1915ꎬ H. Hand. ̄Mazz. 1922       al. 156 ( NAS )ꎻ Changshuꎬ T. Y. Cheo 2178
   (holotype: WU).                                   ( NAS )ꎻ Dongtaiꎬ F. X. Liu 7372 ( NAS )ꎻ
       Synonym: Trapa maximowiczii Korsh.ꎬ Trudy     Kunshanꎬ H. Migo s. n. ( NAS)ꎻ Nanjingꎬ Xuanwu
   Imp. S. ̄Peterburgsk. Bot. Sada 12: 336. 1892.     Lakeꎬ S. L. Chen 24 ( NAS )ꎻ Suzhouꎬ H. Migo
   Type: RUSSIAꎬ without precise localityꎬ Birula s.  s.n. ( NAS)ꎻ Wujiangꎬ F. X. Liu 1555 ( NAS)ꎻ
   n. (holotype: LE!ꎻ photo: P!)ꎻ syn. nov.          Wuxiꎬ Y. H. Law 3029 ( NAS)ꎻ Zhenjiangꎬ East
       Trapa natans L. var. pumila Nakanoꎬ Bot.      China Exped. 2934 ( NAS ). Jiangxi: Guixiꎬ M.
   Mag. ( Tokyo) 77: 166. 1964. Type: VIETNAMꎬ       X. Nie et al. 3848 ( WUK )ꎻ Guangdeꎬ Guangde
   without precise localityꎬ Dec. 1891ꎬ B. Balansa 7421  Bot. Exped. 3163 ( NAS)ꎻ Ji’ anꎬ B. Y. Ding &
   ( holotype: Nakano ’ s Herbariumꎻ isotype: K!)ꎻ   T. Huang 6401 ( HZU )ꎻ Jiujiangꎬ H. Migo s. n.
   syn. nov.                                         (NAS)ꎻ Nanchangꎬ W. H. Wan 770003 ( JXU)ꎻ
                                  野菱  Fig. 3: A      Tangshanꎬ W. H. Wan s. n. ( JXU ). Shanghai:
       Compared with Trapa natansꎬ var. quadricaudata  Hongqiaoꎬ H. Migo s.n. (NAS)ꎻ Minhangꎬ H. Migo

   has relatively smaller fruits and leaves. Leaves of  s. n. ( NAS)ꎬ C. H. Tao 328 ( NAS). Sichuan:
   T. natans var. quadricaudata are rhombicꎬ but com ̄  Huihaiꎬ T. T. Yu 1578 ( PEꎬ IBSC )ꎻ Leiboꎬ Z.
   pressed roundedꎬ incised ̄dentate and thinly dentate at  S. Diao 2504 ( YZU)ꎻ Xichangꎬ S. Y. Chen 10005
   margin in luxuriantly growing status. Flowers are  (NAS). Yunnan: Daliꎬ B. Y. Ding et al. 6248
   whiteꎬ rarely pink in early flowering status. Fruits are  ( HZU)ꎻ Erhaiꎬ Z. S. Diao 2637 (YZU)ꎬ B. Y. Qiu
   frequently bulgedꎻ crowns are smallꎬ beaks conspicu ̄  61150 (KUN)ꎻ Fohaiꎬ C. W. Wang 74239 ( KUNꎬ
   ousꎬ four horns acute and barbellateꎬ sometimes with  PE)ꎬ 77210 (KUNꎬ PE)ꎻ Jianchuanꎬ Jianhu Lakeꎬ
   two lower horns degeneratedꎬ and fresh exocarps   B. Y. Qiu 61192 ( KUN)ꎻ Xiaguanꎬ Erhai Lakeꎬ
   green or red.                                     B. Y. Qiu 60802 ( KUNꎬ IBK )ꎬ 60823 ( KUNꎬ
       Additional collections studied. China. Anhui:  IBK )ꎬ 60882 ( KUNꎬ IBK )ꎻ Xishuangbannaꎬ
   Chuzhouꎬ   anonymous  195   ( NAS )ꎻ   Jinzhaiꎬ   K. M. Feng 20493 ( KUN)ꎻ Yongningꎬ Z. S. Diao
   anonymous 0661 ( NAS )ꎻ Quanjiaoꎬ East China      2577 ( YZU ). Zhejiang: Hangzhouꎬ B. Y. Ding
   Exped. 3507 ( NAS )ꎻ Shuchengꎬ East China         2197 ( HZU)ꎬ 2288 ( HZU)ꎬ 4184 ( HZU)ꎬ S.
   Exped. 4413 ( PEꎬ NAS)ꎻ Wuhuꎬ B. Y. Ding et       Y. Chang 1587 ( NAS )ꎻ Kaihuaꎬ L. Hong 1533
   al. 6216 (HZU)ꎬ 6220 (HZU)ꎻ Xuanchengꎬ anon ̄      (HZU)ꎻ Lishuiꎬ Mt. Nanmingshanꎬ Y. H. He 5981
   ymous 585 (NAS)ꎻ Yixianꎬ B. Y. Ding & T. Huang    (HZU)ꎬ B. Y. Ding 6280 ( HZU )ꎻ Ningboꎬ B.
   6374 ( HZU)ꎬ 6376 ( HZU)ꎻ Yuexiꎬ anonymous        Y. Ding et al. 5786 (HZU)ꎻ Shaoxingꎬ M. Z. Shi &
   2095 ( NAS ). Fujian: without precise localityꎬ   R. Y. Hu 6237 ( HZU)ꎬ 6238 ( HZU)ꎬ B. Y. Ding
   H. H. Chung 3068 (NAS)ꎬ 4579 ( PE). Guizhou:      2190 (HZU)ꎻ Yinxianꎬ M. Z. Shi & R. Y. Hu 6237
   Lipingꎬ S. M. Chang 3066 ( JXU)ꎻ Tianzhuꎬ G.      (HZU)ꎬ B. Y. Ding & K. F. Dong 6429 ( HZU)ꎬ
   X. Ren 770076 (JXU)ꎻ Weiningꎬ S. M. Chang 2322    B. Y. Ding & M. Z. Shi 6210 ( HZU )ꎻ Putuoꎬ
   (IBSC)ꎬ Y. Tsiang 9165 (PE). Hebei: Changpingꎬ    H. Migo s. n. ( NAS )ꎻ Tiantaiꎬ Y. Y. Ho 28230
   X. Y. Liu & Z. S. Zhang s. n. ( PE)ꎻ Fangshanꎬ    ( NAS )ꎬ Zhejiang Bot. Exped. 28406 ( NAS )ꎻ
   Fangshan Exped. 663 ( PE ). Henan: Xuchangꎬ       Wenzhouꎬ B. Y. Ding 4763 ( HZU )ꎻ Wuxingꎬ
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